Joel Sartore Joel Sartore

National Geographic Photographer and Speaker

Keyword: Behavior

Photo: Two captive bred Mexican gray wolves (Canis lupus baileyi) with their mother at the Sedgwick County Zoo.

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Two captive bred Mexican gray wolves (Canis lupus baileyi) with their mother at the Sedgwick County Zoo.

Three king penguins (Aptenodytes patagonicus) on South Georgia Island.

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Three king penguins (Aptenodytes patagonicus) on South Georgia Island.

A king penguin (Aptenodytes patagonicus) rookery from South Georgia Island's St. Andrews Bay. Shown is one of the largest king penguin colonies in the world at 100,000 nesting pairs.

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A king penguin (Aptenodytes patagonicus) rookery from South Georgia Island’s St. Andrews Bay. Shown is one of the largest king penguin colonies in the world at 100,000 nesting pairs.

Photo: A radio-collared lioness and her cub wake up at dusk on their treetop perch in the Ishasha section of Queen Elizabeth National Park.

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A radio-collared lioness and her cub wake up at dusk on their treetop perch in the Ishasha section of Queen Elizabeth National Park.

Photo: The opening spread of the Albertine Rift article, from the November, 2011 issue of National Geographic magazine

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The opening spread of the Albertine Rift article, from the November, 2011 issue of National Geographic magazine

Photo: Approximately 500 wild horses (Equus ferus) at the Triple U Bison Ranch near Fort Pierre, South Dakota.

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Approximately 500 wild horses (Equus ferus) at the Triple U Bison Ranch near Fort Pierre, South Dakota.

Photo: Approximately 500 wild horses (Equus ferus) at the Triple U Bison Ranch near Fort Pierre, South Dakota.

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Approximately 500 wild horses (Equus ferus) at the Triple U Bison Ranch near Fort Pierre, South Dakota.

A brownbelly leaf frog (Phyllomedusa tarsius) collected near Pilalo, Ecuador.

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A brownbelly leaf frog (Phyllomedusa tarsius) collected near Pilalo, Ecuador.

American goldfinch (Carduelis tristis) feed at Waveland Farm near Walton, Nebraska.

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American goldfinch (Carduelis tristis) feed at Waveland Farm near Walton, Nebraska.

American goldfinch (Carduelis tristis) feed at Waveland Farm near Walton, Nebraska.

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American goldfinch (Carduelis tristis) feed at Waveland Farm near Walton, Nebraska.

Photo: A moose (Alces americanus) crosses the Kukpuk River on Alaska's North Slope.

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A moose (Alces americanus) crosses the Kukpuk River on Alaska’s North Slope.

A marsupial frog (Gastrotheca cuencana) at a captive breeding facility in Ecuador. Formerly known as Azuay marsupial frog (Gastrotheca litonedis).

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A marsupial frog (Gastrotheca cuencana) at a captive breeding facility in Ecuador. Formerly known as Azuay marsupial frog (Gastrotheca litonedis).

A meadow vole (Microtus pennsylvanicus pennsylvanicus) from Watertown, New York.

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A meadow vole (Microtus pennsylvanicus pennsylvanicus) from Watertown, New York.

A diamondback water snake (Nerodia rhombifer rhombifer) tries to swallow a catfish at Bayou De View in the Cache River National Wildlife Refuge, Arkansas.

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A diamondback water snake (Nerodia rhombifer rhombifer) tries to swallow a catfish at Bayou De View in the Cache River National Wildlife Refuge, Arkansas.

A diamondback water snake (Nerodia rhombifer rhombifer) tries to swallow a catfish at Bayou De View in the Cache River National Wildlife Refuge, Arkansas.

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A diamondback water snake (Nerodia rhombifer rhombifer) tries to swallow a catfish at Bayou De View in the Cache River National Wildlife Refuge, Arkansas.

A diamondback water snake (Nerodia rhombifer rhombifer) tries to swallow a catfish at Bayou De View in the Cache River National Wildlife Refuge, Arkansas.

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A diamondback water snake (Nerodia rhombifer rhombifer) tries to swallow a catfish at Bayou De View in the Cache River National Wildlife Refuge, Arkansas.

A diamondback water snake (Nerodia rhombifer rhombifer) tries to swallow a catfish at Bayou De View in the Cache River National Wildlife Refuge, Arkansas.

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A diamondback water snake (Nerodia rhombifer rhombifer) tries to swallow a catfish at Bayou De View in the Cache River National Wildlife Refuge, Arkansas.

A diamondback water snake (Nerodia rhombifer rhombifer) tries to swallow a catfish at Bayou De View in the Cache River National Wildlife Refuge, Arkansas.

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A diamondback water snake (Nerodia rhombifer rhombifer) tries to swallow a catfish at Bayou De View in the Cache River National Wildlife Refuge, Arkansas.

A diamondback water snake (Nerodia rhombifer rhombifer) tries to swallow a catfish at Bayou De View in the Cache River National Wildlife Refuge, Arkansas.

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A diamondback water snake (Nerodia rhombifer rhombifer) tries to swallow a catfish at Bayou De View in the Cache River National Wildlife Refuge, Arkansas.

A diamondback water snake (Nerodia rhombifer rhombifer) tries to swallow a catfish at Bayou De View in the Cache River National Wildlife Refuge, Arkansas.

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A diamondback water snake (Nerodia rhombifer rhombifer) tries to swallow a catfish at Bayou De View in the Cache River National Wildlife Refuge, Arkansas.

A diamondback water snake (Nerodia rhombifer rhombifer) tries to swallow a catfish at Bayou De View in the Cache River National Wildlife Refuge, Arkansas.

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A diamondback water snake (Nerodia rhombifer rhombifer) tries to swallow a catfish at Bayou De View in the Cache River National Wildlife Refuge, Arkansas.

A diamondback water snake (Nerodia rhombifer rhombifer) tries to swallow a catfish at Bayou De View in the Cache River National Wildlife Refuge, Arkansas.

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A diamondback water snake (Nerodia rhombifer rhombifer) tries to swallow a catfish at Bayou De View in the Cache River National Wildlife Refuge, Arkansas.

A diamondback water snake (Nerodia rhombifer rhombifer) tries to swallow a catfish at Bayou De View in the Cache River National Wildlife Refuge, Arkansas.

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A diamondback water snake (Nerodia rhombifer rhombifer) tries to swallow a catfish at Bayou De View in the Cache River National Wildlife Refuge, Arkansas.

A Wyoming toad (Anaxyrus baxteri or Bufo baxteri) at the Omaha Zoo. (IUCN: Extinct in the wild; US: Endangered)

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A Wyoming toad (Anaxyrus baxteri or Bufo baxteri) at the Omaha Zoo. (IUCN: Extinct in the wild; US: Endangered)

American goldfinch (Carduelis tristis) feed at Waveland Farm near Walton, Nebraska.

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American goldfinch (Carduelis tristis) feed at Waveland Farm near Walton, Nebraska.

American goldfinch (Carduelis tristis) feed at Waveland Farm near Walton, Nebraska.

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American goldfinch (Carduelis tristis) feed at Waveland Farm near Walton, Nebraska.

American goldfinch (Carduelis tristis) feed at Waveland Farm near Walton, Nebraska.

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American goldfinch (Carduelis tristis) feed at Waveland Farm near Walton, Nebraska.

American goldfinch (Carduelis tristis) feed at Waveland Farm near Walton, Nebraska.

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American goldfinch (Carduelis tristis) feed at Waveland Farm near Walton, Nebraska.

American goldfinch (Carduelis tristis) feed at Waveland Farm near Walton, Nebraska.

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American goldfinch (Carduelis tristis) feed at Waveland Farm near Walton, Nebraska.

American goldfinch (Carduelis tristis) feed at Waveland Farm near Walton, Nebraska.

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American goldfinch (Carduelis tristis) feed at Waveland Farm near Walton, Nebraska.

American goldfinch (Carduelis tristis) feed at Waveland Farm near Walton, Nebraska.

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American goldfinch (Carduelis tristis) feed at Waveland Farm near Walton, Nebraska.

American goldfinch (Carduelis tristis) feed at Waveland Farm near Walton, Nebraska.

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American goldfinch (Carduelis tristis) feed at Waveland Farm near Walton, Nebraska.

American goldfinch (Carduelis tristis) feed at Waveland Farm near Walton, Nebraska.

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American goldfinch (Carduelis tristis) feed at Waveland Farm near Walton, Nebraska.

American goldfinch (Carduelis tristis) feed at Waveland Farm near Walton, Nebraska.

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American goldfinch (Carduelis tristis) feed at Waveland Farm near Walton, Nebraska.

Photo: Julie Jensen Director of Marketing | WVC O: 866.800.7326 | D: 702.443.9249 | E: j.jensen@wvc.org

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