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Joel Sartore Joel Sartore

National Geographic Photographer and Speaker

Keyword: Howler monkey

An endangered (IUCN) and federally threatened female Guatemalan mantled howler (Alouatta pigra) at the Gladys Porter Zoo in Brownsville, Texas.

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An endangered (IUCN) and federally threatened female Guatemalan mantled howler (Alouatta pigra) at the Gladys Porter Zoo in Brownsville, Texas.

Female black howler monkeys (Alouatta caraya) in the Pantanal, Brazil.

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Female black howler monkeys (Alouatta caraya) in the Pantanal, Brazil.

Female black howler monkeys (Alouatta caraya) in the Pantanal, Brazil.

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Female black howler monkeys (Alouatta caraya) in the Pantanal, Brazil.

Photo: A capuchin monkey (Cebus cay) in the Pantanal, Brazil.

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A capuchin monkey (Cebus cay) in the Pantanal, Brazil.

Photo: A capuchin monkey (Cebus cay) jumps from tree to tree in the Pantanal, Brazil.

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A capuchin monkey (Cebus cay) jumps from tree to tree in the Pantanal, Brazil.

Photo: A capuchin monkey (Cebus cay) jumps from tree to tree in the Pantanal, Brazil.

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A capuchin monkey (Cebus cay) jumps from tree to tree in the Pantanal, Brazil.

Photo: A capuchin monkey (Cebus cay) in the Pantanal, Brazil.

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A capuchin monkey (Cebus cay) in the Pantanal, Brazil.

Female black howler monkeys (Alouatta caraya) in the Pantanal, Brazil.

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Female black howler monkeys (Alouatta caraya) in the Pantanal, Brazil.

A close-up of a male black howler monkey (Alouatta caraya) in the Pantanal, Brazil.

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A close-up of a male black howler monkey (Alouatta caraya) in the Pantanal, Brazil.

A female black howler monkey (Alouatta caraya) and her baby in the Pantanal, Brazil.

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A female black howler monkey (Alouatta caraya) and her baby in the Pantanal, Brazil.

A male black howler monkey (Alouatta caraya) in the Pantanal, Brazil.

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A male black howler monkey (Alouatta caraya) in the Pantanal, Brazil.

A male black howler monkey (Alouatta caraya) screams in the Pantanal, Brazil.

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A male black howler monkey (Alouatta caraya) screams in the Pantanal, Brazil.

A male black howler monkey (Alouatta caraya) in the Pantanal, Brazil.

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A male black howler monkey (Alouatta caraya) in the Pantanal, Brazil.

A male black howler monkey (Alouatta caraya) in the Pantanal, Brazil.

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A male black howler monkey (Alouatta caraya) in the Pantanal, Brazil.

A male black howler monkey (Alouatta caraya) in the Pantanal, Brazil.

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A male black howler monkey (Alouatta caraya) in the Pantanal, Brazil.

A male black howler monkey (Alouatta caraya) in the Pantanal, Brazil.

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A male black howler monkey (Alouatta caraya) in the Pantanal, Brazil.

A male black howler monkey (Alouatta caraya) screams in the Pantanal, Brazil.

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A male black howler monkey (Alouatta caraya) screams in the Pantanal, Brazil.

A male black howler monkey (Alouatta caraya) in the Pantanal, Brazil.

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A male black howler monkey (Alouatta caraya) in the Pantanal, Brazil.

A male (black) and female (brown) black howler monkeys (Alouatta caraya) in the Pantanal, Brazil.

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A male (black) and female (brown) black howler monkeys (Alouatta caraya) in the Pantanal, Brazil.

Female black howler monkeys (Alouatta caraya) in the Pantanal, Brazil.

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Female black howler monkeys (Alouatta caraya) in the Pantanal, Brazil.

A female black howler monkey (Alouatta caraya) and her baby in the Pantanal, Brazil.

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A female black howler monkey (Alouatta caraya) and her baby in the Pantanal, Brazil.

A female black howler monkey (Alouatta caraya) and her baby in the Pantanal, Brazil.

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A female black howler monkey (Alouatta caraya) and her baby in the Pantanal, Brazil.

A female black howler monkey (Alouatta caraya) in the Pantanal, Brazil.

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A female black howler monkey (Alouatta caraya) in the Pantanal, Brazil.

Female black howler monkeys (Alouatta caraya) in the Pantanal, Brazil.

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Female black howler monkeys (Alouatta caraya) in the Pantanal, Brazil.

Female black howler monkeys (Alouatta caraya) in the Pantanal, Brazil.

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Female black howler monkeys (Alouatta caraya) in the Pantanal, Brazil.

Female black howler monkeys (Alouatta caraya) in the Pantanal, Brazil.

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Female black howler monkeys (Alouatta caraya) in the Pantanal, Brazil.

A female black howler monkey (Alouatta caraya) and her baby in the Pantanal, Brazil.

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A female black howler monkey (Alouatta caraya) and her baby in the Pantanal, Brazil.

Female black howler monkeys (Alouatta caraya) in the Pantanal, Brazil.

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Female black howler monkeys (Alouatta caraya) in the Pantanal, Brazil.

A female black howler monkey (Alouatta caraya) and her baby in the Pantanal, Brazil.

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A female black howler monkey (Alouatta caraya) and her baby in the Pantanal, Brazil.

A female black howler monkey (Alouatta caraya) and her baby in the Pantanal, Brazil.

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A female black howler monkey (Alouatta caraya) and her baby in the Pantanal, Brazil.

A female black howler monkey (Alouatta caraya) swings from the tree branches in the Pantanal, Brazil.

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A female black howler monkey (Alouatta caraya) swings from the tree branches in the Pantanal, Brazil.

A female black howler monkey (Alouatta caraya) in the Pantanal, Brazil.

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A female black howler monkey (Alouatta caraya) in the Pantanal, Brazil.

Female black howler monkeys (Alouatta caraya) in the Pantanal, Brazil.

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Female black howler monkeys (Alouatta caraya) in the Pantanal, Brazil.

Female black howler monkeys (Alouatta caraya) hang from the trees in the Pantanal, Brazil.

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Female black howler monkeys (Alouatta caraya) hang from the trees in the Pantanal, Brazil.

Female black howler monkeys (Alouatta caraya) in the Pantanal, Brazil.

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Female black howler monkeys (Alouatta caraya) in the Pantanal, Brazil.

Photo: Julie Jensen Director of Marketing | WVC O: 866.800.7326 | D: 702.443.9249 | E: j.jensen@wvc.org

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