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A rancher’s dog expresses displeasure by snarling at photographer, Joel Sartore, on a ranch in Idaho.

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A wild jaguar snarls over her meal in Bolivia’s Madidi National Park.

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A wild jaguar snarls over her meal in Bolivia’s Madidi National Park. The wounds in her shoulder are from a fight with another cat.

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A wild jaguar snarls over her meal in Bolivia’s Madidi National Park. The wounds in her shoulder are from a fight with another cat.

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A Siberian tiger at the Henry Doorly Zoo in Omaha, Nebraska.

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A trained wolf snarls at the camera near Heber City, Utah.

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A trained wolf snarls at the camera near Heber City, Utah.

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A trained wolf snarls at the camera near Heber City, Utah.

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A trained wolf snarls at the camera near Heber City, Utah.

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A trained wolf snarls at the camera near Heber City, Utah.

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A trained wolf snarls at the camera near Heber City, Utah.

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A trained wolf snarls at the camera near Heber City, Utah.

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A trained wolf snarls at the camera near Heber City, Utah.

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A trained wolf snarls at the camera near Heber City, Utah.

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A trained wolf snarls at the camera near Heber City, Utah.

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A trained wolf snarls at the camera near Heber City, Utah.

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A captive wolf at the International Wolf Center near Ely, MN, defends a road-killed deer carcass from the other members of its pack.

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A wolf at the International Wolf Center in Ely, MN, snarls over a road-killed deer, defending its meal from other wolves.

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A wolf at the International Wolf Center in Ely, MN, snarls over a road-killed deer, defending its meal from other wolves.

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A wolf at the International Wolf Center in Ely, MN, snarls over a road-killed deer, defending its meal from other wolves.

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A wolf at the International Wolf Center in Ely, MN, takes a bite of a road-killed deer.

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A sheltie smiles at the camera in a Lincoln, Nebraska neighborhood.

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A sheltie smiles at the camera in a Lincoln, Nebraska neighborhood.

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A sheltie smiles at the camera in a Lincoln, Nebraska neighborhood.

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A sheltie smiles at the camera in a Lincoln, Nebraska neighborhood.

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A sheltie smiles at the camera in a Lincoln, Nebraska neighborhood.

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A sheltie snarls at the camera in a Lincoln, Nebraska neighborhood.

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A dog riding shotgun in a pickup snarls at the photographer.

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A wild jaguar snarls over her meal in Bolivia’s Madidi National Park.

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A wild jaguar snarls over her meal in Bolivia’s Madidi National Park.

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A wild jaguar snarls over her meal in Bolivia’s Madidi National Park.

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A wild jaguar snarls over her meal in Bolivia’s Madidi National Park.

Photo: Julie Jensen Director of Marketing | WVC O: 866.800.7326 | D: 702.443.9249 | E: j.jensen@wvc.org

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